Table 1. Founders and awardees.



Science background

Sergey Brin

Cofounder of Google Inc.

Bachelor's degree in mathematics and computer science from the University of Maryland, College Park and a master's degree in computer science from Stanford University, where he is currently on leave from the PhD program

Priscilla Chan


Bachelor's degree in biology from Harvard University and a medical degree from the University of California, San Francisco

Art Levinson

Chairman of Apple, chairman of Roche's Genentech Inc. unit and member on the board of directors for Roche. Chairman of the board of directors for the Breakthrough Prize in Life Sciences Foundation

Bachelor's degree in molecular biology from the University of Washington and a PhD in biochemical sciences from Princeton University

Yuri Milner

Founder of the internet company Group. Member of the board of directors for the Breakthrough Prize in Life Sciences Foundation

Advanced degree in theoretical physics from Lomonsov Moscow State University and subsequently conducted research at the Institute of Physics at the Russian Academy of Sciences

Anne Wojcicki

Cofounder of privately held personal genetics company 23andMe Inc. Member of the board of directors for the Breakthrough Prize in Life Sciences Foundation

Bachelor's degree in biology from Yale University

Mark Zuckerberg

Founder, chairman and CEO of Facebook Inc. Member of the board of directors for The Breakthrough Prize in Life Sciences Foundation

Studied computer science at Harvard



Research focus

Cornelia Bargmann

Professor and head of the Lulu and Anthony Wang Laboratory of Neural Circuits and Behavior at The Rockefeller University and a Howard Hughes Medical Institute (HHMI) investigator

Genetics of neural circuits and behavior, and synaptic guidepost molecules

David Botstein

Director of the Lewis-Sigler Institute for Integrative Genomics at Princeton University and professor of genomics at Princeton

Linkage mapping of Mendelian disease in humans using DNA polymorphisms

Lewis Cantley

Professor and director of the Cancer Center at Weill Cornell Medical College and New York-Presbyterian Hospital

Phosphoinositide 3-kinase (PI3K) and its role in cancer metabolism

Hans Clevers

Professor of molecular genetics at the Hubrecht Institute and president of the Royal Netherlands Academy of Arts and Sciences

Wingless-type MMTV integration site (WNT) signaling in tissue stem cells and cancer

Napoleone Ferrara

Professor of pathology and senior deputy director for basic sciences at the University of California, San Diego Moores Cancer Center

Mechanisms of angiogenesis that lead to therapies for cancer and eye diseases

Titia de Lange

Professor, head of the Laboratory of Cell Biology and Genetics and director of the Anderson Center for Cancer Research at The Rockefeller University

Telomeres-illuminating how they protect chromosome ends and their role in genome instability in cancer

Eric Lander

President and founding director of the Broad Institute of MIT and Harvard, professor of biology at the Massachusetts Institute of Technology (MIT) and professor of systems biology at Harvard Medical School

General principles for identifying human disease genes and enabling their application to medicine through the creation and analysis of genetic, physical and sequence maps of the human genome

Charles Sawyers

Chair of human oncology and the pathogenesis program at Memorial Sloan-Kettering Cancer Center and an HHMI investigator

Cancer genes and targeted therapy

Bert Vogelstein

Director of the Ludwig Institute for Cancer Research, professor of oncology and pathology at The Sidney Kimmel Comprehensive Cancer Center at The Johns Hopkins University School of Medicine and an HHMI investigator

Cancer genomics and tumor suppressor genes

Robert Weinberg

Professor for cancer research at MIT, director of MIT's Ludwig Center for Molecular Oncology and a member of the Whitehead Institute for Biomedical Research

Characterization of human cancer genes

Shinya Yamanaka

Director of the Center for iPS Cell Research and Application at Kyoto University and senior investigator at The J. David Gladstone Institutes

Induced pluripotent stem cells